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Bertula crucialis Felder & Rogenhofer comb. rev.
     Bertula crucialis Felder & Rogenhofer, 1874, Reise öst. Fregatte Novara, Zool 2 (Lepid.): pl. 120: 37.

Bertula crucialis
Figure 99

The dark brown wings are fasciated and marked by straight sharp white lines, the forewing with straight antemedial and postmedial fasciae and a narrower, slightly angled submarginal. There is a central longitudinal white line associated with the CuA system, an angular ‘8’-shaped reniform in white, and further longitudinal white lines near the costa. The hindwing is similar over its distal half, and has a broader white wedge extending basad from the margin near M1. The male labial palps are upcurved with a slender third segment that has a small tuft of scales subapically on its dorsal side.

Taxonomic note. Poole (1989) transferred this and the next species to Foveades Bethune-Baker (type species aroensis Bethune-Baker, New Guinea) without comment, together with bisectalis Wileman (Taiwan). It is better to retain them in a broad concept of Bertula until such a time as this and allied genera can be revised in totality. The new species that follow have some general resemblance in facies, but differ in other features as discussed for each.

Geographical range. Sundaland.

Habitat preference. Apart from one specimen from lower montane forest at 1000m, the species has been recorded most frequently in lowland forest, including heath forest. Chey (1994) recorded five from softwood plantations and secondary forest near Brumas in the lowlands of Sabah.

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