Miscellaneous Genera VI
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Ilyrgis Walker

Type species: echephurealis Walker, Sri Lanka.

This genus,
Parilyrgis Bethune-Baker and the next two genera are characterised by small, rather delicate mauvish grey species with angled or bifalcate forewings and bipectinate male antennae.

Ilyrgis species have the ground of the wings relatively uniform, with fine, irregular white antemedial, postmedial and submarginal fasciae on the forewing, these tending to broaden at the costa. The hindwings are angled marginally at CuA2, with more diffuse pale postmedial and submarginal fasciae similarly angled. The labial palps are upcurved, the second segment long, erect, the third less than half its length and angled slightly forwards.

The male abdomen has an eighth segment of the framed corematous type, though rather weakly expressed. The genitalia have the uncus somewhat bulbous towards the apex. The valves are simple, tongue-like (the type species) or with slight steps in the centre of the costal and saccular margins, the valve narrowing beyond where they occur (costinotata). The aedeagus is short, slightly curved. with an ovate vesica, large and extensively spined in costinotata, but moderate and much more sparsely spined, the spines shorter, in the type species.

In the female genitalia (type species) the ostium is between the seventh and eighth segments. The ductus bursae is as long as the seventh segment, slender, unsclerotised. The corpus bursae is pyriform, densely invested with short spines over its distal half.

The genus consists of the two species below and
capnosia Hampson (New Guinea), but the placement in it of other species from New Guinea, Africa and Madagascar listed by Poole (1989) needs checking.

The only Australian representative amongst the four genera,
Goniocraspedon mistura Swinhoe, is assigned to the Hypeninae in Nielsen et al. (1996), though Nye (1975), Inoue et al. (1982) and Poole (1989) retained them in the Ophiderinae. The angling of the hindwing submarginal, the simple valve structure, the spining in the vesica of some males and in the ductus bursae of some females in the group is perhaps more typical of Herminiinae than Hypeninae. Indeed, Forbes (1954) regarded the very similar New World genus Redectis Nye (= Dercetis Grote, praeocc.) as one of the more primitive of the Herminiinae, a placement retained by Poole (1989). Ectogonia butleri Leech (see below) is noted to prefer dried leaves of its hosts, a further herminiine characteristic.

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