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Myrioblephara Warren

Type species: rubrifusa Warren, N.E. Himalaya.

Synonym: Prorhinia Warren (type species pingasoides Warren (N.E. Himalaya) syn. n.

This and the next six genera are probably closely related, sharing a number of morphological characters. Sato (1984a) included two of them in his Aethalura McDunnough group. The genus-group names Necyopa Walker and Ectropidia Warren have priority over Diplurodes Warren but taxa referable to them have all been assigned to Diplurodes in the past (Holloway, 1976). Holloway (1991) made a preliminary segregation of species between Diplurodes, Ectropidia and Necyopa: in the treatment here these are restored as full genera. Three new genera are described to embrace species excluded by the stricter definition of Diplurodes, and a further one for a species misplaced in Ectropis Hübner.

In all except Necyopa (where they are bipectinate) the male antennae are fasciculate. In most genera there are pairs of intersegmental coremata on the male abdomen, unusual in the Boanniini. The signum in the female genitalia, when present, is a transverse flange with a smoothly curved margin. The ovipositor is elongate, extensile in some species but not to the extent seen in Ectropis Hübner or Cleora Curtis. In the male, abdominal sternite 3 lacks a setal comb, and the forewing is without a fovea in Diplurodes, Ectropidia and two of the new genera.

In Myrioblephara the male coremata tend to be reduced to a pair between segments 6 and 7, though the type species has rudimentary ones between 7 and 8; the type species of Prorhinia has weak pairs in both positions. In the male genitalia the uncus is short, broad, often bifid. The gnathus is strong. The juxta is usually an elongate, straplike sclerotisation. The valve costa is either broad, uniformly and strongly setose, apically protrusive (both generic type species) or shortened and somewhat separated as a distinct arm (two of the new Bornean species). The sacculus usually bears a rod-like process, apically setose and directed towards the apex of the valve: from the base of this a narrow band of sclerotisation runs across the valve to near the base of the costa.

These genitalic structures show diverse modification amongst the species, and the definition and extent of the genus needs further study. A swarm of species in New Guinea (e.g. M. flexilinea Warren, M. mollis Warren, M. muscosa Warren, M. subtrita Warren) lacks coremata in the male abdomen and has genitalia that do not conform very closely to the definition above. In facies they are similar to the two commoner Bornean species. Sato (1993) transferred a number of Himalayan species to Myrioblephara.

Sato (1984a) noted that the larva of a Japanese species had setae D1 on A2-5 and A8 raised basally, creating small dorsolateral angles to the outline. The colour is an irregularly mottled and banded ligneous brown (Sugi, 1987). The host-plant was Quercus (Fagaceae).

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