Type species: lugubris Walker.
Synonyms: Metorthocheilus Hampson (type species emarginata Hampson,
cited erroneously as subobscurata Walker by Fletcher (1979)) syn. n.; Paradirades
Warren (type species assimilis Warren, New Guinea).
The above synonymy brings together genera where the male has a
characteristic double fovea at the base of the forewing, straddling, and divided
by, the fold marking the position of vein CuP: the basal part of vein CuA is
somewhat sinuous just anterior to the fovea. Also combined with the genus are
two further species that lack this fovea but have a sinuous base to vein CuA and
other diagnostic features of the male genitalia described below. All have rather
narrow wings, the hindwings usually with two straight, parallel fasciae rather
than the typically angled ones seen in most epiplemines. The male antennae are
densely unipectinate, approaching the serrate, lamellate condition of other
The male genitalia have a pair of spines, probably modified setae, at
the distal end of the sacculus, the major diagnostic feature. The costa may also
terminate in a spine or spur (not in lugubris and assimilis). The
uncus is entire, narrow (emarginata), broad (lugubris and assimilis)
or bifid (the two additional species). The gnathus is present but weak in
the first two but strong, divided in the last. The first two have a small
hair-pencil at the base of the valve sacculus, absent in the two additional
species, though these have a sparsely setose, digitate process basally on the
interior of each valve.
The ovipositor lobes of the female are usually squarish or triangular.
The bursa has two small stellate signa opposite a much larger one (assimilis:
Fig 203), two large stellate signa (emarginata) or two longitudinal
bands of scattered small spines (the additional species).
All known species are mentioned above or in the section following,
except for C. fulvilunata Warren (New Guinea). One has been reared from
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