Olene inclusa Walker
1856, List Specimens lepid. Insects Colln Br. Mus., 7:1737.
1856, List Specimens lepid. Insects Colln Br. Mus., 7:1724.
 1858-9, Cat. Lepid. Insects Mus. E. Ind. Co., 2: 340.
1862, J. Linn. Soc. (Zool.), 6:140.
invaria demaculata Strand,
1915, Gross-Schmett. Erde, 10: 293.
sensu Holloway, 1976: 51.
Diagnosis. The forewings are much broader than in congeners, with the discal lunule
enlarged, rather oblique, more conspicuous in forms with a uniformly dark brown
forewing. The rarer form with pale blotches as in the typical form of mendosa
has the broad central area extending beyond the postmedial rather than being
bounded by it. Females also have a broader forewing, with the blacker zone being
more transverse and postmedial in position.
Taxonomic note. There are no genitalic differences between males of inclusa and invaria,
hence the two taxa are treated as synonyms. Though invaria has page
priority over inclusa, those authors (e.g. van Eecke, 1928) who have
regarded them as conspecific have treated invaria as the junior synonym.
This must stand on the ‘first reviser’ principle. O. dalbergiae Moore
comb. n. (N. India) is also closely related, having very similar male
genitalia, as has O. hypersceles Collenette comb. n. (Sulawesi).
Geographical range. Sundaland, Philippines, Sulawesi, Sumbawa.
Habitat preference. The majority of records are from a range of lowland
forest types, but the species also extends to as high as 1670m.
Biology. Moore (in Horsfield & Moore, 1859 ) illustrated the larva in
Java. It is dark brown with dorsolateral white bands and a grey ventral surface.
The secondary setae are dark, some plumose at their tips. The dorsal brushes are
rufous brown, paler at the base. There are hair-pencils front and rear as in
mendosa, but lateral ones on A2 are absent. Gardner (1938) indicated that the
larva of dalbergiae also lacked the lateral tufts of mendosa.
The host range is not as large as in O. mendosa but includes the
following (Moore; Pholboon, 1965; Kalshoven, 1981; Kuroko & Lewvanich, 1993;
unpublished IIE, H.S. Barlow and FRIM records): Annona (Annonaceae); Averrhoa
(Averrhoaceae); Durio (Bombacaceae); Shorea (Dipterocarpaceae);
(Euphorbiaceae); Leea (Leeaceae); Pelargonium (Geraniaceae); Acacia,
Crotalaria, Derris, Erythrina, Mucuna (Leguminosae); Ficus (Moraceae);
Musa (Musaceae); Calyptranthes, Eugenia (Myrtaceae); Rosa (Rosaceae);
Citrus (Rutaceae); Theobroma (Sterculiaceae); Muntingia (Tiliaceae);
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