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Olene inclusa Walker  
Dasychira inclusa Walker, 1856, List Specimens lepid. Insects Colln Br. Mus., 7:1737.
Repena invaria Walker, 1856, List Specimens lepid. Insects Colln Br. Mus., 7:1724.
Dasychira asvata Moore, [1860] 1858-9, Cat. Lepid. Insects Mus. E. Ind. Co., 2: 340.
Thelde patula Walker, 1862, J. Linn. Soc. (Zool.), 6:140.
Dasychira invaria demaculata Strand, 1915, Gross-Schmett. Erde, 10: 293.
Dasychira hypersceles Collenette sensu Holloway, 1976: 51.

Olene inclusa

Olene inclusa

Olene inclusa

The forewings are much broader than in congeners, with the discal lunule enlarged, rather oblique, more conspicuous in forms with a uniformly dark brown forewing. The rarer form with pale blotches as in the typical form of mendosa has the broad central area extending beyond the postmedial rather than being bounded by it. Females also have a broader forewing, with the blacker zone being more transverse and postmedial in position.

Taxonomic note. There are no genitalic differences between males of inclusa and invaria, hence the two taxa are treated as synonyms. Though invaria has page priority over inclusa, those authors (e.g. van Eecke, 1928) who have regarded them as conspecific have treated invaria as the junior synonym. This must stand on the ‘first reviser’ principle. O. dalbergiae Moore comb. n. (N. India) is also closely related, having very similar male genitalia, as has O. hypersceles Collenette comb. n. (Sulawesi).

Geographical range. Sundaland, Philippines, Sulawesi, Sumbawa.

Habitat preference. The majority of records are from a range of lowland forest types, but the species also extends to as high as 1670m.

Biology. Moore (in Horsfield & Moore, 1859 [1860]) illustrated the larva in Java. It is dark brown with dorsolateral white bands and a grey ventral surface. The secondary setae are dark, some plumose at their tips. The dorsal brushes are rufous brown, paler at the base. There are hair-pencils front and rear as in mendosa, but lateral ones on A2 are absent. Gardner (1938) indicated that the larva of dalbergiae also lacked the lateral tufts of mendosa.

The host range is not as large as in O. mendosa but includes the following (Moore; Pholboon, 1965; Kalshoven, 1981; Kuroko & Lewvanich, 1993; unpublished IIE, H.S. Barlow and FRIM records): Annona (Annonaceae); Averrhoa (Averrhoaceae); Durio (Bombacaceae); Shorea (Dipterocarpaceae); Ricinus (Euphorbiaceae); Leea (Leeaceae); Pelargonium (Geraniaceae); Acacia, Arachis, Crotalaria, Derris, Erythrina, Mucuna (Leguminosae); Ficus (Moraceae); Musa (Musaceae); Calyptranthes, Eugenia (Myrtaceae); Rosa (Rosaceae); Citrus (Rutaceae); Theobroma (Sterculiaceae); Muntingia (Tiliaceae); Congea (Verbenaceae).

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