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Etanna Walker

Type species: basalis Walker, Borneo.

Andrapha Walker (type species basalis); Apothripa Hampson (type species iphida Swinhoe, India); Bagistana Walker (type species rudis Walker = breviuscula Walker, Borneo) syn. n.; Clettharra Walker (type species valida Walker = breviuscula, Borneo) syn. n.; Nanaguna Walker (type species breviuscula Walker, Borneo) syn. n.; Tamusida Walker (type species vittalis Walker, Borneo) syn. n.; Thrypticodes Lucas (type species xyloglypta Lucas = breviuscula, Australia) syn. n.

This genus and its new synonyms are brought together on shared features of forewing venation and male abdominal features. The included species also show a wide variety of male secondary sexual characters, and several show sexual dimorphism. Many also have white or pale marginal flecks at around one-third from the tornus of the forewing. The venation feature is in the radial sector branching system which is (R2 ((R3, R4) R5)), the branching off of R5 being unusually distal to that of R2; there is no areole. The hindwing venation is quadrifid, with M3 and CuA1 stalked, except in the type species of Etanna where it is trifid, and the male has the wing strongly bilobed. In E. breviuscula comb. n. and E. teleoleuca Prout comb. n. the male hindwing has a sharp subbasal angle to the costa, the costal zone has tufts of dark brown scales, and there is a dark scale pencil within the cell just posterior to these.

Other secondary sexual characters are found in the male abdomen. Lateral coremata (three pairs) are found between segments 4, 5, 6 and 7 in basalis, breviuscula and teleoleuca. Coremata are also found in association with the seventh and eighth segments in some other species. There are no tymbal organs. The eighth segment is short, the apodemes of both sclerites short, broad, widely spaced; in some species (e.g. E. brunnea Hampson comb. n.) the eighth sternite becomes enlarged and expanded to direct the genitalia into a more upwardly facing orientation. The genitalia have an uncus much as in
Characoma Walker, simple, tapering from a broad base, but the tegumen, though elongate and sometimes expanded slightly on each side, or produced on each side of the uncus (breviuscula and allies), is not more expanded at the junction with the vinculum and does not overlap it. The valves are diagnostically reduced in size and are usually smaller than the processes that bear the clump of dark scales, and this is usually globular, rarely blade-like as in Characoma. The saccus is broad and well developed.

The female genitalia have the ovipositor lobes acute. The ductus is very long and slender. The corpus bursae is usually spherical and with general spining rather than a signum, though the spining may vary in development across the corpus bursae.

The genus is restricted to the Indo-Australian tropics but contains several species that are widespread thoughout this area. The biology of several species is known, as discussed individually below.

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