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Meganola Dyar

Type species: conspicua Dyar, U.S.A.

This genus-group name was used by Poole (1989) to include all taxa previously assigned (e.g. Hampson, 1900) to
Roeselia Hübner (see Franclemont, 1960; Poole, 1989; Holloway & Miller, 1995) except a few such as Sarbena and Proneca that he treated as distinct. As discussed on p. 14, these genera (though not typical Meganola (Hampson, 1900)) and Manoba (a record of loose stacking) and a small number of others include records of head capsule stacking in the larvae. All except Evonima have quadrifine hindwing venation with M3 and CuA1 stalked (almost to the margin in M. conspicua). The forewing venation is as in Ctenane (see above), with all radial sector veins present (with a small areole in some species, including conspicua). Holloway & Miller (1995) indicated this concept was paraphyletic and associated with it taxa such as Manoba and Rhynchopalpus that were excluded from their strict concept of Nola. However, these taxa have reduced forewing venation and trifine hindwing venation that, in combination with features of the male abdomen, enable them to be separated from Meganola, leaving a smaller paraphyletic assemblage that has, by and large, complete venation, bipectinate male antennae, rather plesiomorphic male genitalia and the more advanced ‘invaginated signum’ type of female genitalia.

A number of Bornean species are here placed under this more restricted but probably paraphyletic concept of Meganola, as no strong classificatory signal was detected in features of the male and female genitalia that would permit new generic groupings to be recognised.

However, the first four species treated and the ones to which they are related share a slight sexual dimorphism in the hindwing: uniform grey in females; grading paler towards the dorsum in males.

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