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Beara nubiferella Walker
Beara nubiferella Walker, 1866, List Specimens lepid. Insects Colln Br. Mus., 35: 1704.
milbradti Kobes, 1997: 81, syn. n.


Beara nubiferella

All three Bornean species are very similar in facies and are only reliably distinguished on genitalic characters. This and the next species have similarly dark scales on the male coremata, and a costal thorn to the valve and a cornutus in the aedeagus vesica, but the thorn is angled basad in nubiferella and the cornutus is small, slender. The female has a similarly thickened, broad ductus and bursa in each, but nubiferella has a distinctive narrow tubular pocket at the apex.

Taxonomic note. Holloway (1982) underestimated the complexity of the situation and focused on male characters. Kobes (1997) noted sympatry of cornuta Holloway with the species he described as milbradti. This latter now proves on dissection of the holotype female of nubiferella, to be a synonym thereof. The commoner species in Java and Bali that Holloway took to be nubiferella (slides 10429, 10450, 10451, 17339) is in fact undescribed and closely related to
tortriciformis Strand, though with a more rounded valve costa. A further member of this group is B. simplex Warren from Adonara, east of Flores in the Lesser Sundas. The Sulawesi variant of cornuta mentioned by Holloway (1982) also appears to be a distinct species, having a long, straight cornutus in the aedeagus vesica and a much obtuser, less thorn-like projection to the valve costa. A Sulawesi female dissected (slide 17335) has the ductus and bursa unthickened, but is not as in the tortriciformis Strand group.

Geographical range. Java, Sumatra, Borneo.

Habitat preference. This is possibly the commonest of the Bornean species, but all are rare. Five specimens have been noted, all from the lowlands: one from Kretam on the coast of Sabah, two from hill dipterocarp forest at the Danum Valley Field Centre and two from alluvial dipterocarp forest near the foot of G. Mulu.

Biology. Robinson et al. (2001) note host records from Ceiba (Bombacaceae) and Tectona (Verbenaceae; Mathur, 1942), but, given the taxonomic problems, these records should perhaps be considered at the generic level only.

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