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Daddala lucilla Butler
Lacera sublineata Walker, 1865, List Specimens lepid. Insects Colln Br. Mus., 33: 1030.]
Sypna lucilla Butler, 1881, Trans. ent. Soc. London, 1881: 206.
Sypna obscurata Butler, 1881, Trans. ent. Soc. London, 1881: 207.
Daddala lucilla Butler; Holloway, 1976: 31; Kobes, 1985: 63.

Daddala lucilla Daddala lucilla Daddala lucilla

Males of this species have strongly bipectinate antennae. The forewing pattern is very variable, in both sexes, and is much more irregular than in all congeners except the last one listed below.

Taxonomic note. An attempt was made to relate the complex of three species recognised by Kobes (1985) in Sumatra to the situation in Borneo. Five males that appeared to correspond to the range of underside variation identified by Kobes were dissected. All had very similar genitalia, the aedeagus having a of small spicules just distal to the anellus at the basal end of the origin of the vesica and a further one within the small distal expansion of the spines on the column of the aedeagus itself. Two Himalayan males lacked the spicules but had more obtuse spines (e.g. as in the holotype of obscurata Butler, illustrated by Berio & Fletcher).

A senior synonym for
lucilla may exist in sublineata Walker, noted to be a Daddala species by A. Zilli (pers. comm.), though listed under its original genus, Lacera, by Poole (1989). The male type (Cambodia; in UM, Oxford) definitely falls within the lucilla complex, but its status will be unclear until the findings of Kobes are tested further.

The specimen illustrated for Nepal as lucilla by Haruta (1993: Plate 51, fig. 13) matches the holotype of the Himalayan D. renisigna Moore in facies precisely. However, Berio & Fletcher (1958) regarded renisigna as distinct from lucilla and a senior synonym of D. expressa Prout (Sumatra), which has distinctive pale hindwings very different from those of renisigna. The male genitalia of expressa are also distinctive as described by Kobes (1985), who followed Berio & Fletcher in treating it as a synonym of renisigna. The holotype of renisigna has lost the distal part of its abdomen, so a comparison of its genitalia has yet to be undertaken. It is likely that this synonymy is erroneous, that renisigna is referable to lucilla, and that expressa is a valid species endemic to Sumatra, but no Himalayan specimen matching renisigna so precisely has been located in BMNH.

Geographical range. Himalaya, Taiwan, Japan, Burma, Thailand (VK), Sumatra, Borneo, Java, Bali, Sulawesi.

Habitat preference. The species is frequent in montane forest from 1000m to 2600m.

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