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Rusicada nigritarsis Walker comb. rev.
Rusicada nigritarsis Walker, [1858] 1857, List Specimens lepid. Insects Colln Br. Mus., 15: 1006.
Anomis albipuncta Snellen, 1880, Tijdschr. Ent. 23: 76.
Gonitis xanthochroa Butler, 1886, Trans. ent. Soc. Lond., 1886: 409.
Anomis ambonensis Swinhoe, 1920, Ann. Mag. nat. Hist. (9), 5: 257, syn. n.
Anomis irene Prout, 1929, Bull. Hill Mus. Witley, 3: 11, syn. n.

Rusicada nigritarsis
(Sri Lanka)

. This and all the species except the last have the typical orange-red forewings, but nigritarsis and revocans Walker are distinguished by the reniform where the posterior lobe is smaller than the anterior one, rather than equal to it or larger. The average direction of the zig-zag anterior part of the postmedial is oblique to the costa in nigritarsis and more or less at right-angles to it in revocans. In the male genitalia the robust juxta bifurcations extend almost to the apex of the valve and are curved in nigritarsis, much shorter and straight in revocans. In both species the setose projections from the anellar attachment to the aedeagus are asymmetrical and the valve coremata have the setal bases entirely pale. In the females the lobes of the lamella postvaginalis are broader at the base than they are long. Those of revocans have an angular process between the lobes of the lamella postvaginalis, and there is a diverticulum at the base of the ductus bursae.

Taxonomic note. The subspecies (see below) are separated on facies differences: nigritarsis is uniform greyish red; albipuncta is a dark, dull, brick-red; xanthochroa is dark as in albipuncta but mottled with broad areas of paler colour. New Caledonian material is somewhat intermediate between albipuncta and xanthochroa in characters.

Geographical range. Sri Lanka, India, Hainan, Taiwan, Borneo, Java; Sulawesi, Moluccas, Queensland, New Caledonia (ssp. albipuncta); Solomons, Vanuatu, Fiji, Samoa, Tonga (ssp. xanthochroa).

Habitat preference. The general distribution suggests a preference for seasonally dry areas of the tropics. In New Caledonia (Holloway, 1979) and Fiji (Robinson, 1975) the species was taken mostly in open, disturbed or secondary habitats. In Borneo a female was recorded in an area of coastal and secondary vegetation at Seria in Brunei.

Biology. Moore (1884-1887) described two forms of larva which he attributed to nigritarsis. The green form may well be referable to albitibia (see below) which is sympatric in Sri Lanka, and the other form, olive grey with fine longitudinal white lines, each segment dotted black, the head yellowish, may possibly be true nigritarsis.

Host-plants recorded (Robinson, 1975; Holloway, 1979; Robinson
et al., 02001) are Hibiscus and Urena (Malvaceae).

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