View Image Gallery of Tribe Ophiusini

Artena Walker

Type species: submira Walker, India.

This and the next two genera may be related, but it is difficult to determine the polarity of various characters in relation to their states in the Achaea / Parallelia complex of genera discussed next. The species tend to be very robust in body, particularly the thorax. The forewings have simpler, more linear forewing fasciation on a much more uniform ground, especially the submarginal fascia which is only broken in some Ophiusa Ochsenheimer species. The reniform is more strongly developed than in the other generic group, particularly in Thyas Hübner (see p. 44). The male genitalia lack coremata on the valves (or these are much reduced). The larvae are elongate semi-loopers, longitudinally finely striate in pale and dark brown, with Artena and Thyas showing development of an elliptical or circular mark dorsally on A5. The pupa lacks a bloom. Host plants are drawn from several families, but there is a preponderance of records from Combretaceae and Myrtaceae, with Artena only known from the former and Ophiusa Guenée having a higher frequency of Myrtaceae records.

Particular features of Artena are a characteristic forewing facies usually with oblique antemedials and postmedials, and a submarginal very close to the margin, cutting off a paler, often rather mauve marginal zone. The reniform is less prominent than in the other two genera. The hindwing has a medial blue band that stops well short of the anterior and posterior margins. This is shared with two Thyas species.

The male abdomen has the eighth sternite twice as broad as it is long and as the tergite, shallowly cleft on its posterior margin, and with the anterior margin rather convex. The tergite may have short, broad, well separated apodemes. The genitalia have the uncus associated with a scaphium, and often with a short subbasal spur dorsally. The valves are deeply based over the vinculum which is slightly longer than the tegumen. Their processes usually show some degree of bilateral asymmetry, and consist of a long one arising from the base of the costa and a series traversing the valve from this point to the apex of the sacculus. The distal part of the valve is reduced, tongue-like. The juxta is of the inverted ‘V’ type. The aedeagus is broad at the base and variably flexed more distally. The vesica tends to be relatively small, though the diverticula can be long and narrow.

In the female of the type species, the ostium is within a complex, rather ring-like sterigma in the position of the seventh sternite. The ductus is very short, and the corpus bursae is elongate, irregular in width, angled in places. It is very finely scobinate throughout. The ductus seminalis arises from a small, tapering appendix bursae near the base of the corpus bursae.

The genus is most diverse in the Oriental tropics but two species extend further east, with a third,
A. velutina Prout, restricted to the Papuan Subregion. Artena angulata Roepke is endemic to Sulawesi.

The larva of the type species was described by Bell (MS), who stated it to be indistinguishable in colour, shape and markings from that of
A. dotata Fabricius as described below but also similar to that of Thyas Hübner and Ophiusa Ochsenheimer. The pupa is also similar, and the host plants were all from the Combretaceae as for dotata: Combretum, Getonia, Quisqualis and Terminalia.

<<Back >>Forward <<Return to Content Page

Copyright © Southdene Sdn. Bhd. All rights reserved.