This tribe consists of those genera recognised as a natural group by
Holloway (1984b), with the addition of Celerena Walker (Holloway,
1993), Derambila Walker and allied genera and Ozola Walker (= Desmobathra
Meyrick), as discussed below.
The group is cosmopolitan and almost entirely tropical and subtropical.
Included genera not represented in Borneo are:
New World: Dolerophyle Warren, Dolichoneura Warren, Entogonia Warren, Leptoctenopsis
Warren, Ophiogramma Hübner, Pycnoneura Warren;
Africa: Antozola Herbulot, Apatadelpha Prout, Brachytrite Swinhoe,
Panagropsis Warren, (see also Conolophia Warren below);
Australasia: Caledophia Holloway, Callipotnia Warren, Encryphia Turner, Physetostege
Diagnostic features are found particularly in the male abdomen, notably
in the structure of the tegumen in which the characteristic loops thickening
each side have migrated, looping ventrally below the articulation with the
vinculum leaving a central, elongated column of thickening to support the uncus,
with the lacuna within the lateral loops much reduced, or even absent as in some
species of Ozola. This condition of the tegumen can be seen most clearly
in Figs 40, 48, 50, 69 and 75. The vinculum is normal and coremata are absent.
The abdomen itself is usually very slender, elongated, with a pair of
longitudinally elongated patches of setae on the third sternite. The elongation
of these, when present, is another diagnostic feature (Figs 50, 51). In
the group of genera recognised by Holloway (1984b) there is a recurrent feature
on the eighth segment: a pair of curved, finely corrugate, sausage-shaped zones
laterally (Fig 35). This is seen in Conolophia, Noreia Walker, Alex
Walker and Foveabathra Gen. n. in the Bornean fauna. In
several genera there are also complex structures at the junction of the third
and fourth sternites: it is seen in three of the four genera just mentioned, Alex
being the exception.
The male antennae are usually ciliate, sometimes narrowly bipectinate.
The female genitalia offer no tribal level characteristics. The
ornamentation of the bursa shows great variety, e.g.: general (Noreia)
or localised (Derambila) scobination; longitudinal rows of
spines (Ozola; a crown-like central ring of broad spines (Conolophia,
Callipotnia); a small, scobinate signum (Encryphia); a
longitudinal scobinate band (Celerena); immaculate (Alex, with
an appendix; Foveabathra).
The ansa of the tympanum is usually simple, tapering, thought to be the
plesomorphic condition by Cook & Scoble (1992), sometimes with a slight
Seven genera of the group occur in Borneo.
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