TRIBE LITHOSIINI
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Tigrioides Butler

Type species: alterna Walker, Australia.

Synonym: Parapelosia Bethune-Baker (type species grisescens Bethune­Baker, New Guinea).

Hampson (1900) included the three Bornean species listed below in his concept of Lexis Wallengren which appears to have been based on an erroneous identification of the S. African type species. The species he included now rest by default in Tigrioides, placed by him as a synonym of Lexis. The synonymy above is as in Nielsen et al. (1996). The current assemblage is certainly not a natural grouping, and comments on nitens Walker and allies may be found in Holloway (1979), where they were combined as a natural group tentatively in Eilema Hübner (see Eilema Hübner); nitens is still placed in Tigrioides by Nielsen et al.

Tigrioides alterna has typically lithosiine divided valves in the male genitalia, with a ridge running from the centre of the dorsal part towards the saccular part. The small aedeagus has a single, massive cornutus in the vesica. The female has a massively thickened and flocculent ductus bursae that is as long as the pyriform bursa, central to which are two scobinate patches; the ovipositor lobes are acute. The male of typical Parapelosia has more slender valves lacking a ridge between the two components; the aedeagus is more slender and lacks a cornutus. The female has a very short ductus that is evenly sclerotised, leading into a longer, broader neck section of the corpus bursae, the broader part of which is distally bilobed; the scobinate patches of the signa are reduced, and the ovipositor lobes are square.

The four Bornean species currently placed in Tigrioides share none of these features. The placement of sabulosalis Walker is discussed below. The next two species share general facies similarities such as black spots in the thorax, and both have a spined zone at the apex of the aedeagus which is relatively much larger than in either of the generic type species. In both species the vesica is large with a multiply-spined cornutus and, in leucanioides Walker, a second single one. The females of both have a corpus bursae that is large and irregularly bilobed, a large distal lobe arising beyond a smaller lateral one. Thus the species could be congeneric, but their placement needs further study. The fourth species is new, but closely related to aurantiaca Hampson from the Philippines. Its features are also discussed below.

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