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Cozola Walker

Type species: leucospila Walker, Sulawesi.

Synonym: Pronygmia Toxopeus (type species xanthopera Hampson, India, Sumatra) syn. n.

The facies of the forewings of Cozola species is consistent in form, though variable in colour, the bulk of the species in Sulawesi being black and white, the Sundanian and Philippines ones more brown and yellow. All have a row of irregularly sized (some very reduced) white patches in the spaces of the distal margin of the forewing. The postmedial and antemedial of the forewing are distinctly zig-zag or lunulate, with the medial zone often a paler tone than the rest of the wing. The radial system has lost a vein, probably R5, with M1 showing a slight stalk with Rs.

The male genitalia are also diagnostic, and tymbals are present in the abdomen. The uncus is well developed, but without striking modifications. The valves are simple, rather tongue-like, with long setae around the costal and distal margins and on a lobe or process on the sacculus. The saccus is moderate, narrow, with slight interior swellings on each side at its base. The aedeagus is a simple tube, the vesica unadorned.

The female genitalia (xanthopera) have rather square ovipositor lobes, the eighth segment is a simple ring with fine setae moderately densely all over. The ductus is short, narrow, the bursa flimsy, elongate-oval, without a signum.

The biology of the Bornean species is described below.

As well as the species below and the generic type species, Cozola already includes a large complex of species in Sulawesi. The following additional species are congeneric: C. hapala Collenette (Peninsular Malaysia, Sumatra, Java, Bali) comb. n.; C. austriaca Semper comb. n. (Philippines); C. geometrica
Semper comb. n. (Philippines); C. acroptera Collenette comb. n. (Sulawesi);C. menadoensis Collenette comb. n. (Sulawesi); C. paloe Collenette comb. n. (Sulawesi).

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