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Quadricalcarifera viridimaculata Matsumura  
Quadricalcarifera viridimaculata Matsumura, 1922, Zool. Mag., Tokyo 34: 521.
Quadricalcarifera bioculata Kiriakoff, 1967:  42, syn. n.; Kiriakoff 1968: 120;Holloway 1976: 55,
        fig. 364.

Quadricalcarifera viridimaculata

Quadricalcarifera viridimaculata

The well defined white spots on the green forewing of the male are distinctive; in trioculata (below) they are more blurred, with a distal dentate process from the reniform. The subapical bar on the hindwing in both sexes is broader than in trioculata and the margin of the wing is less evenly rounded. In the female the greenish-brown patches of the forewing are more clearly defined, there are no central submarginal spots and the black lines across the costal zone are less oblique.

Taxonomic notes. Though externally this species resembles the next, the male genitalia suggest a relationship with eusebia Kiriakoff (Sumatra) and allies.

The species was identified from the illustration of the holotype (Taiwan) in Sugi (1979). Female specimens from the N.E. Himalaya may have been confused in the past with fasciata Moore, described from the male only, which is strikingly different from the male of viridimaculata but tends towards the female in having a broad white marginal band to the forewing. Possibly the taxon hasegawai, described from Malaya by Nakamura (1976), will prove to be a synonym of viridimaculata, but the species was not illustrated by its author.

Geographical range. N.E. Himalaya, China, Taiwan, Sundaland, Mindanao.

Habitat preference. This is a strictly montane species, taken from 1050 - 2600 m on G. Kinabalu and frequently at 1618 m on Bukit Retak in Brunei. The lowest record is from 1000 m on G. Mulu. The species thus ranges from the lower to upper montane forest zones.

Biology. Sevastopulo described a larva which produced an adult that he attributed to fasciata (Sevastopulo 1938 - 47(3)). As it was a female it could have been viridimaculata misidentified but the specimen has not been located. The head of the larva is bluish green, the body green with a pale yellow collar; there are fawn blotches dorsally on each segment; the spiracles and legs are red, the prolegs green. The pupa is subterranean in an earthen cocoon. The host was Quercus (Fagaceae). Gardner (1946) recorded Pterocarya (Juglandaceae) as a host for fasciata but again this identification cannot be checked from voucher specimens.

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