TRIBE SCOLIOPTERYGINI
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Savara latimargo Walker 
Gonitis latimargo Walker, [1858] 1857, List Specimens lepid. Insects Colln Br. Mus., 13: 1002.
Gonitis brunnea Moore, 1882, Descr. new Indian lepid. Insects Colln W.S. Atkinson, 2: 153, syn. n.
Zethes variabilis Pagenstecher, 1888, Jb. nassau Ver. Naturk., 43: 54, syn. n.
Savara longipectinata Prout, 1922, Bull. Hill Mus. Witley, 1: 241, syn. n.
Savara catortha Hampson, 1926, Descr. Gen. Spec. Noctuinae, p. 344, syn. n.

Savara latimargo
(Philippines)
Savara latimargo
(Philippines)


Diagnosis
. This is a somewhat larger and browner species than its congeners, slightly paler on the upperside and more so on the underside where the postmedial fasciae are diagnostically darker, and lunulate in each space. The submarginal at the forewing apex is an arc of black dashes, usually with white highlighting on its distal side. The forewing postmedial above is straighter than in other species, particularly in the female and in material from Sundaland eastwards. The male abdomen has a distinctly square invagination to the anterior margin of the eighth sternite, and the valve of the genitalia is expanded apically to appear squarely bilobed. In the female, the base of the ductus is sclerotised and tubular over a greater extent than in congeners, with a strong, tongue-shaped excavation in the ventral margin of the ostium.

Taxonomic note. The type material in BMNH of both latimargo and brunnea is from India, but is lacking abdomina. Specimens from Burma (slides 19344 and 19345) are taken to represent the species. These specimens share the diagnostic features described above with the Australasian variabilis Pagenstecher. This was placed in its genus of original description by Poole (1989), but was assigned to Savara by Edwards in Nielsen et al. (1996), with longipectinata and catortha as synonyms. The female (slide 18738) included in type material of longipectinata is in fact referable to pratti (see above). The larval description by Bell below was attributed to brunnea, and it is evident from Australasian material labelled as brunnea in his collection in BMNH that his identification was probably correct, though voucher specimens for the rearing have not been located.

Geographical range. India, Burma, Borneo, Philippines, Sulawesi, Moluccas to Queensland and Solomons.

Habitat preference
. The only Bornean specimen seen is a female from Samarinda in the lowlands of Kalimantan.

Biology. The larva in India was reared by Bell (MS). The prolegs on A3 are considerably reduced, those on A4 slightly so. The anal claspers splay out behind. Early instars are elongate, transparent white, with obscure darker spots laterally on each segment. Later instars are olive-green, marked with pale purple and with the setae arising from black dots. The mature larva is lightish brown, each black setal dot ringed with white. There is a pale dorsal line that has short lateral branches near the anterior margin of each segment; there are obscure paler patches outside this line, and more obscure patching and lineation on the flanks.

The eggs are pale bluish green, flattened, dome-shaped, with 50 shallow ribs linked by numerous fine cross-rays. Pupation is in a cell between two leaves of the host plant. The pupa lacks a powdery bloom. The larvae were reared on
Grewia (Tiliaceae), but Bell suspected it was not the natural host. The species has been reared from Sterculia (Sterculiaceae) in New Guinea (Miller et al., unpublished).

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