TRIBE OPHIUSINI
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Ophiusa Ochsenheimer

Type species: tirhaca Cramer, South Africa.

Synonyms: Hemachra Sodoffsky (unnecessary replacement name for Ophiusa); Meropis Hübner (type species auricularis Hübner, Europe = tirhaca); Ophiogenes R.L. (type species tirhaca).

This genus is possibly related to the previous two as discussed on p. 40. However, the species all have much narrower forewings and more irregular and, in some species, broken fasciation. The submarginal tends to be more prominent subapically, and the reniform and orbicular are often conspicuous, but without the variability seen in Thyas. The ground colour, particularly of the hindwings and abdomen, is often yellow, more occasionally fawn.

In the male genitalia the uncus has a distinct basal spur or crest dorsally. There is much bilateral asymmetry in the tegumen, juxta and valves, the costal processes of the latter tending to be short, spur-like. In circumferens Guenée, triphaenoides Walker and allies the exterior of the distal part of the valve is extensively but shallowly corematous. The basal part of the aedeagus is usually broadened and upturned, and the vesica is highly convolute.

In the female genitalia (Fig 20, tirhaca), the ostium is set well to the anterior of the seventh sternite, mostly posterior to which is a variously complex and bilaterally asymmetric sterigma. The ductus is convolute, sometimes with a lateral lobe, and the corpus bursae is pyriform and generally scobinate.

Bell (MS) described the larva of the type species. The prolegs on A4 are half the size of those on A5 and A6, and those on A3 are even further reduced. The colour is lichen-grey to smoky black, banded longitudinally, particularly by a blackish dorsolateral line, and stippled along the bands with rufous dots. There are blackish lines and dots dorsally on A5, but the circular dorsal spot seen in Thyas and Artena (see below) appears to be absent, as in the species following where the larva is known. There is a transverse, tumid ridge on A8 that bears the dorsolateral tubercles and is lined black posteriorly.

The pupa lacks a powdery bloom. The type species is widespread in the Old World tropics and subtropics, including Australasia (Holloway, 1979; Common, 1990) and occurs in Sumatra (Kobes, 1985), but has not so far been recorded from Borneo, where the related O. tirhacoides sp. n. occurs. Several other species are widespread in the Indo-Australian tropics and occur in Borneo, and species of the more localised kenricki Bethune-Baker group are found in the Australasian tropics (Robinson, 1975; Holloway, 1979).

Host records for the type species throughout its range (Holloway, 1979; Common, 1990; Robinson
et al., 2001) are Mangifera, Pistacia, Rhus (Anacardiaceae), Cistus (Cistaceae), Combretum (Combretaceae), Shorea (Dipterocarpaceae), Erica (Ericaceae), Pelargonium (Geraniaceae), unspecified Loranthaceae, Eucalyptus, Leptospermum, Myrtus, Psidium (Myrtaceae) and Rosa (Rosaceae). Robinson et al. noted various fruit crop records from Zhang (1994) but these probably refer to adult feeding. The genus as a whole, as will be seen below, shows some concentration of records in the Myrtaceae.

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