species: caradrinoides Guenée, Java.
(type species marginata Moore, 1884 (nec Moore, 1882), Sri Lanka = xanthostola
(type species quadrilineata Walker, Borneo); Eudragana
(type species limbata Butler, Christmas I.); Lacibisa
(type species bifaria Walker, Borneo); Sillophora Warren
(type species bimaculata Warren = bimaculata Snellen,
Sulawesi) praeocc.; Trichoptya Warren
(type species sejuncta Walker).
highly distinctive facies and characteristics of the male abdomen, yet a number
of species has been attached to the genus erroneously and others have been
described in totally unrelated genera.
was redescribed by Kobes (1984, Heterocera Sumatrana 2: 30) as Mecodina
sumatrana Kobes; Mecodina proves to be a better generic placement for this
species (see p. 321). Kobes also described Allocosmia sugii Kobes
from Sumatra in the same paper. The species megastigmata
(Indian Subregion), transferred to Bocula by Poole (1989), is closely related and
should also be placed in Allocosmia (annotations in BMNH by W. Speidel and
the author) under its replacement name Imosca
& Sasaki (2001) comb. n. Poole
(1989) also transferred the taxon padanga Swinhoe to Bocula,
but this proved to be a careine nolid that is now placed in Didigua
(Kobes, 1997; Holloway, 2003).
antennae are fasciculate, sometimes strongly so. The male hindwings can be
variously modified with androconial scales and tufts, and the hindtibia can also
bear a massive tuft of scales or be virtually unmodified. The forewing facies
typifies the genus, consisting of fine, oblique, sometimes curved or obscure
dark antemedial, medial and postmedial fasciae that usually run more or less
parallel to each other. The submarginal may be regularly curved, irregularly so
or subcostally notched, and frequently defines a wholly or partially blackened
marginal area. The reniform is represented by a black dot of various sizes but
this can be obscure; it occurs between the medial and the postmedial, usually
closer to the former.
male abdomen, the eighth sternite is elongate and massively bilobed both
anteriorly and posteriorly, the posterior lobes being invested with hairs and
sometimes subsidiary processes. The tergite is either of similar length and
paddle-like, the anterior handle part of the paddle possibly representing the
elongation and fusion of the two apodemes seen in the second group, as it has an
elongate central lacuna. In the second group (to which the name Borsippa
might be assigned at subgeneric level) the tergite is shorter, triangular with a
distal angle, anteriorly with a pair of broad, widely spaced apodemes that
represent the other angles of the triangle. There are extensive setal areas
lateral to this triangle. The genitalia have a rather weak but elongate,
digitate uncus that (typical group) may have a fringe of setae around its apex.
The valves are typically paddle-like with various small basal and saccular
processes and coremata, or (Borsippa) narrower, lacking a corema and with more extensive
basally directed hairs at the apex and a lobe with setae at two thirds on the
ventral margin. The juxta in both groups is a robust inverted ‘V’, the
components of which do not entirely fuse when they meet centrally.
female genitalia, the ostium is broad and situated well within the eighth
segment. It tapers into a short to moderate ductus. The corpus bursae is
elongate and may be generally scobinate or have a ridge-like signum. The seventh
sternite is somewhat reduced and posteriorly rounded. In subgenus Borsippa
ostium is slightly more anterior, between the seventh and eighth segments, and
is not as broad. The seventh sternite is relatively more reduced and has a
distinctly bilobed posterior margin. The ductus is also very short (though the
neck of the bursa may be long and as narrow), and the bursa is variable in
shape, pyriform (e.g. in divergens
or with a long neck.
differs from the typical group in facies, usually having extensive black in the
forewing marginal area; these species are treated from xanthostola
genus is predominantly Oriental, but extends east to New Guinea the Solomons and
Australia and has several species in Africa.
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