Miscellaneous Genera II
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Bocula Guenée

Type species: caradrinoides Guenée, Java.

Synonyms: Aramuna Moore (type species marginata Moore, 1884 (nec Moore, 1882), Sri Lanka = xanthostola Hampson); Borsippa Walker (type species quadrilineata Walker, Borneo); Eudragana Butler (type species limbata Butler, Christmas I.); Lacibisa Walker (type species bifaria Walker, Borneo); Sillophora Warren (type species bimaculata Warren = bimaculata Snellen, Sulawesi) praeocc.; Trichoptya Warren (type species sejuncta Walker).

Species of Bocula have highly distinctive facies and characteristics of the male abdomen, yet a number of species has been attached to the genus erroneously and others have been described in totally unrelated genera.

B. poaphiloides Walker was redescribed by Kobes (1984, Heterocera Sumatrana 2: 30) as Mecodina sumatrana Kobes; Mecodina proves to be a better generic placement for this species (see p. 321). Kobes also described Allocosmia sugii Kobes from Sumatra in the same paper. The species megastigmata Hampson (Indian Subregion), transferred to Bocula by Poole (1989), is closely related and should also be placed in Allocosmia (annotations in BMNH by W. Speidel and the author) under its replacement name Imosca Sugi & Sasaki (2001) comb. n. Poole (1989) also transferred the taxon padanga Swinhoe to Bocula, but this proved to be a careine nolid that is now placed in Didigua Walker (Kobes, 1997; Holloway, 2003).

The male antennae are fasciculate, sometimes strongly so. The male hindwings can be variously modified with androconial scales and tufts, and the hindtibia can also bear a massive tuft of scales or be virtually unmodified. The forewing facies typifies the genus, consisting of fine, oblique, sometimes curved or obscure dark antemedial, medial and postmedial fasciae that usually run more or less parallel to each other. The submarginal may be regularly curved, irregularly so or subcostally notched, and frequently defines a wholly or partially blackened marginal area. The reniform is represented by a black dot of various sizes but this can be obscure; it occurs between the medial and the postmedial, usually closer to the former.

In the male abdomen, the eighth sternite is elongate and massively bilobed both anteriorly and posteriorly, the posterior lobes being invested with hairs and sometimes subsidiary processes. The tergite is either of similar length and paddle-like, the anterior handle part of the paddle possibly representing the elongation and fusion of the two apodemes seen in the second group, as it has an elongate central lacuna. In the second group (to which the name
Borsippa Walker might be assigned at subgeneric level) the tergite is shorter, triangular with a distal angle, anteriorly with a pair of broad, widely spaced apodemes that represent the other angles of the triangle. There are extensive setal areas lateral to this triangle. The genitalia have a rather weak but elongate, digitate uncus that (typical group) may have a fringe of setae around its apex. The valves are typically paddle-like with various small basal and saccular processes and coremata, or (Borsippa) narrower, lacking a corema and with more extensive basally directed hairs at the apex and a lobe with setae at two thirds on the ventral margin. The juxta in both groups is a robust inverted ‘V’, the components of which do not entirely fuse when they meet centrally.

In the female genitalia, the ostium is broad and situated well within the eighth segment. It tapers into a short to moderate ductus. The corpus bursae is elongate and may be generally scobinate or have a ridge-like signum. The seventh sternite is somewhat reduced and posteriorly rounded. In subgenus Borsippa the ostium is slightly more anterior, between the seventh and eighth segments, and is not as broad. The seventh sternite is relatively more reduced and has a distinctly bilobed posterior margin. The ductus is also very short (though the neck of the bursa may be long and as narrow), and the bursa is variable in shape, pyriform (e.g. in divergens Prout) or with a long neck.

Borsippa group differs from the typical group in facies, usually having extensive black in the forewing marginal area; these species are treated from xanthostola Hampson onwards.

The genus is predominantly Oriental, but extends east to New Guinea the Solomons and Australia and has several species in Africa.

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