Type species: polyclealis Walker.
This new genus brings together species from both traditional Chloroclystis
and Gymnoscelis that have generally grey wings, crossed by angular,
blackish postmedials that often have areas of white immediately distal to them.
Males do not have any obvious secondary sexual modifications. They are further
defined by features of the male and female abdomen. The apodemes of the second
tergite are moderate, extending across half the segment and narrow.
In the male the octavals are vestigial or absent. The valves do not have
strong scent pencils and are short, basally deep, tapering to the apex, and
slightly upcurved. The vesica is generally scobinate but also has one to many
larger cornuti. Where there are many, the aedeagus is often strongly reflexed,
such that both base and apex are directed distally.
The female has a short ductus, usually sclerotised, with the bursa set
on a neck of variable length dependent on the development of the aedeagus of the
male. The more extreme this is, the larger and more spiralled is the neck of the
bursa. Both neck and bulb are invested extensively with a dense array of
As well as the species discussed in the descriptions of Bornean taxa,
the genus also includes G. immixtaria Walker comb. n. (= ablechra Turner
syn. n.; Sri Lanka, Chagos Is., Queensland, Fiji (the unidentified Gymnoscelis
of Robinson (1975)), G. expedita Prout comb. n. (Peninsular Malaysia)
and G. acygonia Swinhoe comb. n. (N.E. Himalaya). A similar,
highly reflexed aedeagus is seen in "Chloroclystis” invisibilis Warren
(New Guinea) and "C." macroaedeagus Holloway (New Caledonia) that
have facies more typical of some Gymnoscelis species, and also in "C.”
azumai Inoue (Ryukyu Is.) where the female genitalia are consistent with
those of Glaucoclystis, but the strong octavals and bulbous sacculus of
the male valve are more as in the other Pacific species. “C.” spinosa Inoue
(Ryukyu Is.) also has female genitalia characteristic of Glaucoclystis.
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