View Image Gallery of Tribe Eupitheciini

Chloroclystis Hübner

Type species: coronata Hübner, = v-ata Haworth, Europe.

Synonyms: Aetheolepis Warren (type species papillosa Warren, N.E. Himalaya) praeocc. Chloroplintha Warren (type species velutina Warren, Sulawesi); Dyserga Petersen (type species coronata); Gymnopera Warren (type species rubroviridis Warren, N.E. Himalaya); Oligoclystia Bastelberger (type species blanda Bastelberger, Taiwan); Sesquiptera Warren (type species imnequata Warren, N.E. Himalaya); Simotricha Warren (type species lucinda Butler, Japan; ssp. of v-ata).

The concept of Chloroclystis (e.g. as in Holloway, 1976) has until recently, embraced a large number of species of diverse morphology, united only through possession of two pairs of spurs on the male hind-tibia relative to one in Gymnoscelis Mabille. Examination of the genitalia has indicated this concept to be untenable. An attempt is made here to identify more natural groupings of species.

True Chloroclystis have a number of diagnostic features: the forewing postmedial is distally dentate anteriorly, but often fades away posteriorly; there is sexual dimorphism in the hindwing, that of the male tending to be reduced, with modified scaling dorsally, that of the female being more typically fasciated; similar modified scaling may occur on the underside of the forewing where it overlaps the hindwing; the cross vein between M1 and M2 of the male hindwing is strongly displaced basad relative to that of the female in some species (but not v-ata); the male abdomen lacks octavals but there are paired corematous pouches associated with tergite 7 and sometimes 5 in the Bornean species and some others; there are no coremata associated with the genitalia, the uncus is broadly based, but itself very short; the labides are large, broad, falcate; the valves have few setae and are broad over the basal half, much narrower distally; the aedeagus vesica sometimes has a single large cornutus; the bursa of the female is enlarged, with no definitive signum, but extensive fields of fine, delicate spines. The labial palps are short.

The larva of the type species was described and illustrated by Carter & Hargreaves (1986). It is relatively robust, variably coloured in green, brown or grey, with darker triangular dorsal markings and lateral diagonals on each segment.

The host range is very wide (Allan, 1949), and the larva is a flower-feeder. Pupation is in the soil (Carter & Hargreaves, 1986).

In addition to the generic type species listed above and the Bornean species following, the genus in its strict sense includes: C. luciana Prout (N. India), C. conversa Warren (N.E. Himalaya), C. neoconversa Inoue (Ryukyu Is.), C. olivata Warren (S. India, Sri Lanka), C. nudifunda Warren (Peninsular Malaysia), C. inaequata Warren (N.E. Himalaya, Java, Bali); C. mira Prout (Luzon), C. velutina Warren (Sulawesi), C. palmaria Prout (Sumatra, Java, Peninsular Malaysia), C. analyta Prout (Sumatra), and C. permixta Prout (Java).

The Bornean species are somewhat segregated, though overlapping, with altitude, though all are montane. The only endemic has the highest altitude range.

A number of other genus-group names in addition to those revived for Bornean species in the text following have been subordinated to Chloroclystis in the past (but are not included as synonyms by Nielsen, Edwards & Rangsi), and probably merit revival also. Preliminary investigation of genitalic characters has indicated that Polysphalia Warren (type species cristigera Warren, New Guinea) Gen. rev. and Ptychotheca Warren (type species pallidivirens Warren, New Guinea) Gen. rev. are distinct, possibly congeneric, with male genitalia similar to those of Bosara Walker (see below). Dasimatia Warren (type species subusta Warren, Sulawesi) Gen. rev. is also distinct, with no clear affinities indicated by the female genitalia dissected. Mesocolpia Warren (type species subcomosa Warren, Sao Thomé) Gen. rev. is also distinctive and might be applied to a large number of African 'Chloroclystis' such as consobrina Warren, dexiphyma Prout, lita Prout, marmorata Warren, nanula Mabille, peremptata Walker, protrusata Warren and tumefacta Prout, though subcomosa itself has somewhat modified wings in the male, not seen in the other species.

<<Back >>Forward <<Return to Contents page

Copyright © Southdene Sdn. Bhd. All rights reserved.